Peralec a Doly u Luze, dve lokality cenomanske flory ze Zeleznych hor

In this contribution, plant mega- and meso- fossils from the Peruc-Korycany Formation of the eastern part of the Bohemian Cretaceous Basin are described. They come from localities near the villages of Perálec (temporary outcrop of a sewage treatment plant west of the village) and Doly u Luže permanent outcrop in steep bank of a creek running through the village) in the Pardubice Region. The plant fossils occur in sandy mudstone layers (the Peruc Member) cropping out in basal parts of the Peruc-Korycany Formation. The plant fossils are numerous compressions of entire-margined, lanceolate leaves of angiosperms. Narrowly lanceolate leaves of “Grevillea” constans Velenovský with narrow attenuate apex and cuneate base, and lanceolate leaves of Dicotylophyllum sp. with broadly cuneate base and attenuate apex are both common there. Trilobed dentate leaves with cuneate base of Araliaephyllum formosum (Heer) Greguš et J. Kvaček are rare. Plant mesofossils obtained by bulk maceration of sediment samples provided abundant megaspores of Sellaginaceae Verrucitriletes cf. dubius (Dijkstra) Potonié, and Isoetaceae Paxillitriletes cf. midas (Dijkstra) Hall et Nicolson. Angiosperm mesofossil remains were represented by rarely occurring small elongated-ovoid seeds of Spermatites sp., and larger circular seeds described as Carpolithes sp. Numerous mesofossils of Mauldinia cf. bohemica Eklund et J. Kvaček are represented by bi-lobed structures with three typically circular scars, located basally. They are interpreted as cladodia, a part of compound inflorescence / infructescence. They are associated with leaves of “Grevillea” constans. Fossil charcoalified angiosperm wood is assigned to lauraceous taxon Paraphyllanthoxylon cf. marylandense Herendeen. It shows a diffuse-porous wood with indistinct growth rings.
Based on the analysis of preliminary fossil record, particularly on the presence of Mauldinia and megaspores of lycophytes, a reconstruction of the palaeoenvironment of both localities is suggested to have been a river flood plain (vegetation dominated by Mauldinia) running through coastal lowlands (abundant megaspores of lycophytes).

References

Batten, D. J. - Colin, J.-P. - Néraudeau, D. (2010): Megaspores from mid Cretaceous deposits in western France and their biostratigraphic and palaeoenvironmental significance. - Rev. Palaeobot. Palynol. 161, 151-167.

Batten, D. J. - Zavattieri, A. M. (1996): Re-examination of seed cuticles from Cretaceous deposits in West Greenland. - Cretac. Res. 17, 69-714.

Čech, S. (2011): Palaeogeography and stratigraphy of the Bohemian Cretaceous Basin (Czech Republic) - an overview. - Geol. Výzk. Mor. Slez. 2011/1, 18-21.

Čech, S. - Klein, V. - Kříž, J. - Valečka, J. (1980): Revision of the Upper Cretaceous stratigraphy of the Bohemian Cretaceous Basin. - Věst. Ústř. Úst. geol. 55, 277-296.

Dijkstra, S. J. (1951): Wealden megaspores and some other fossils from the Aachenian (Senonian) in South Limburg, Netherlands. - Meded. Geol. Sticht. Nieuwe Ser. 3, 19-32.

Drinnan, A. N. - Crane, P. R. - Friis, E. M. - Pedersen, K. R. (1990): Lauraceous flowers from the Potomac Group (mid-Cretaceous) of eastern North America. - Bot. Gaz. 151, 3, 370-384.

Eklund, H. - Kvaček, J. (1998): Lauraceous inflorescences and flowers from the Cenomanian of Bohemia (Czech Republic, Central Europe). - Int. J. Plant Sci., 159, 668-686.

Friis, E. M. - Pedersen, K. R. -Crne, P. R. (1999): Early Angiosperm Diversification: The Diversity of Pollen Associated with Angiosperm Reproductive Structures in Early Cretaceous Floras from Portugal. - Ann. Miss. Bot. Gdn 86, 2, 259-296.

Frumin, S. - Eklund, H. - Friis, E. M. (2004): Mauldinia hirsuta sp. nov., a new member of the extinct genus Mauldinia (Lauraceae) from the Late Cretaceous (Cenomanian-Turonian) of Kazakhstan. - Int. J. Plant Sci. 165, 5, 883-895.View article

Gandelová, L. - Šlezingerová, J. (2014): Stavba dřeva. - 187 str. Mendel. univ. v Brně. Brno.

Gomez, G. - Daviero-Gomez, V. - Coiffard, C. - Martín-Closas, C. - Dilcher, D. L. (2015): Montsechia, an ancient aquatic angiosperm. - PNAS 12 (35), 10985-10988.

Greguš, J. - Kvaček, J. (2015): Revision of Cenomanian flora from the Maletín Sandstone. - Acta Mus. Nat. Pragae, Ser. B, Hist. Nat. 71(3-4), 315-364.

Gryc, V. - Vavrčík, H. - Sakala, J. (2009): Cenomanian angiosperm wood from the Bohemian Cretaceous Basin, Czech Republic. - IAWA, 30, 319-329.View article

Herendeen, P. S. (1991): Lauraceous wood from the mid-Cretaceous Potomac group of eastern North America: Paraphyllanthoxylon marylandense sp. nov. - Rev. Palaeobot. Palynol. 69, 277-290.

IAWA Committee (1989): IAWA List of microscopic features for hardwood identification. International. - Int. Assoc. Wood Anatomists Bull. 10, 219-332 (New series).

InsideWood (2004-onwards): Published on the Internet: http://insidewood.lib.ncsu.edu/search?31 [přístupné 28. února, 2018].

Knobloch, E. (1984a): Megasporen aus der Kreide von Mitteleuropa. - Sbor. geol. věd. Paleont. 26, 157-195.

Knobloch, E. (1984b): Megasporen aus der Kreide (Oberconiak-Santon) der Bohrung Volfartice (Nordbühmen). - Čas. Mineral. Geol. 29, 2, 155-165.

Kutluk, H. - Hills, L. V. (2017): Megaspores from the Upper Cretaceous (Campanian) Horshoe Canyon Formation of south- central Alberta, Canada, with a review of the genera Costatheca and Spermatites. - Palynology 41, 31-71.View article

Kvaček, J. (1992a): Leaf and fruit compressions from the Bohemian Cenomanian. In: Kovar-Eder, J., ed.: Palaeovegetational development in Europe and regions relevant to its palaeofloristic evolution, Proceedings of the Pan-European Palaeobotanical Conference, Vienna, 19-23 September 1991, 301-304. - Mus. Nat. History, Vienna.

Kvaček, J. (1992b): A new Cenomanian floral locality from Praha [Nová lokalita cenomanské flóry]. - Čas. Nár. muz., Ř. přírodověd. 158(1-4), 42.

Kvaček, J. (2000): Frenelopsis alata and its microsporangiate and ovuliferous reproductive structures from the Cenomanian of Bohemia (Czech Republic, Central Europe). - Rev. Palaeobot. Palynol., 112, 51-78.

Kvaček, J. - Doyle, J. A. - Endress, P. K. - Daviero-Gomez, V. - Gomez, B. - Tekleva, M. (2016): Pseudoasterophyllites cretaceus from the Cenomanian (Cretaceous) of the Czech Republic: A possible link between Chloranthaceae and Ceratophyllum. - Taxon 65, 6, 1345-1373.View article

Manum, S. B. (1996): Clitellate cocoons. In: Jansonius, J. - McGregor, D.C., ed.: Palynology: Principles and Applications. Volume 1, 361-364. - Amer. Assoc. Stratigraph. Palynologists Fnd, College Station, Texas.

Manum, S. B., Bose, M. N., Sawyer, R. T. (1991): Clitellate cocoons in freshwater deposits since the Triassic. - Zoologica Scripta, 20:347-366. View article

McLoughlin, S. - Bomfleur, B. - Mörs, T. - Reguero, M. A. (2016): Fossil clitellate annelid cocoons and their microbiological inclusions from the Eocene of Seymour Island. - Antarctica. Palaeont. Electronica 19, 1-27.

Oakley, D. - Falcon-Lang, H. J. - Gasson, P. (2009): Morphometric analysis of some Cretaceous angiosperm woods and their extant structural and phylogenetic analogues: implications for systematics. - Rev. Palaeobot. Palynol. 157, 375-390.

Pacltová, B. (1977): Cretaceous angiosperms of Bohemia - Central Europe. - Bot. Rev. 43, 128-142.

Uličný, D. - Kvaček, J. - Svobodová, M. - Špičáková, L. (1997): High-frequence sea-level fluctuations and plant habitats in Cenomanian fluvial to estuarine successions: Pecínov quarry, Bohemia. - Palaeogeogr. Palaeoclimat. Palaeoecology136, 165-197.View article

Vajskebrová,M. - Gürtlerová, P., ed. (1998): Databáze významných geologických lokalit [databáze online]. [citováno 2018-25-02]. Dostupné z URL http:// lokality.geology.cz. - Čes. geol. služba. Praha,

Viehofen, A. - Hartkopf-Fröder, Ch. - Friis, E. M. (2008): Inflorescences and Flowers of Mauldinia angustiloba sp. nov. (Lauraceae) from Middle Cretaceous Karst Infillings in the Rhenish Massif, Germany. - Int. J. Plant Sci., 169, 7, 871-889.

Wheeler, E., A. (1986): Vessels per square millimetre or vessel groups per square millimetre? - Int. Assoc. Wood Anatomists Bull. 7, 73-74 (New series).

Wheeler, E. A. (2011): InsideWood - a web resource for hardwood anatomy. - Int. Assoc. Wood Anatomists J. 32, 199-212.